Thanks all for the (sensible) ideas. I spent an evening fiddling with spoke tensions, replaced one cracked nipple, got everything true again and all spokes pinging at about the same note when plucked, but still have the noises this morning. I suspect it is just one spoke, maybe at the spoke-nipple-rim interface, but I've got no nearer locating it.
Once the noise starts, it is mostly one louder pinging followed by 2 or 3 fainter ones occurring every few seconds but not in a regular timing. There are then occasional pings/clunks at a different note but it is mostly as above.
Must be on the wheels as it happens with and without pedaling, and only really gets going at about 20mph. For info they are Mavic Open Pro rims.